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In addition, autoantigens (25, 26), urate crystals (27, 28), low-density lipoprotein (oxLDL) (29, 30), cholesterol (31), high-mobility group box 1 protein (HMGB1) (32, 33), pro-inflammatory cytokines (e.g., IL-1β (34), IL-8 (35), and tumor necrosis factor-α (TNF-α) (36)) can also stimulate the formation of NETs, which in turn can cause the development of a variety of diseases

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In cases of SLC31A1 deficiency [46], copper uptake can also occur through CTR2, which is encoded by SLC31A2 , as a compensatory mechanism [47]