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The following chemicals were obtained from Sigma-Aldrich: N-acetylcysteine (NAC), Tiron, Trolox, N-ethyl maleimide (NEM) and 4′,6-Diamidino-2-phenylindole (DAPI)

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Then the aged mice were intraperitoneally injected with FOXO4-DRI (5 mg/kg) in PBS or PBS alone every other day for three administrations